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Acknowledgements We gratefully acknowledge funding from The Scottish Government's Rural and Environment Science and Analytical Services Division (RESAS). We are grateful to the Bioservices Division, Moredun Research Institute, for expert care and assistance with animals. We also thank the staff of Parasitology Department at Moredun Research Institute for their technical assistance with the controlled efficacy test, and Steve Paterson for his assistance in analysing initial life history data. ; Peer reviewed ; Publisher PDF